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Paramagnetic relaxation enhancement (PRE) experiments were performed using a Bruker 600 MHz NMR spectrometer with a cryogenic probe (GIST, Gwangju)

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-oxidation initiation: After acyl-CoA enters the mitochondrial matrix, it is gradually degraded through the -oxidation pathway, generating 1 molecule of FADH, 1 molecule of NADH, and 1 molecule of Acetyl-CoA (acetyl-COA) in each cycle