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The Rac1 activator, phorbol 12-myristate 13-acetate (PMA), significantly increased the protein expression levels of MMP-2, -3, -9, and -19, whereas rhein and the Rac1 inhibitor, NSC23766, produced opposite effects
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However, the current largest body of evidence, in the field of metabolic adaptation of cancer cells, deals with the flexible utilization of glutamine and glucose as the main sources of energy
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